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- ArticleGomot L, Bride J.C R Seances Soc Biol Fil. 1976 Oct;170(3):559-83.Azetidine-2-carboxylic acid, a proline analogue which depresses collagen deposition in extracellular spaces, stops uropygial invaginations morphogenesis and epithelial buds branching in vitro. The results show that uninterrupted collagen synthesis is necessary for preen gland morphogenesis.