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- ArticleKhovrychev MP, Korolev PN, Bulgakova VG.Mikrobiologiia. 1978 May-Jun;47(3):442-5.The effect of copper ions and unfavourable pH values of the medium on the incorporation of labelled precursors of protein and RNA was studied in Candida utilis. Specific inhibition of protein synthesis by copper ions and alkaline conditions was found. No specific inhibition of protein or RNA was detected at low pH values of the medium.