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- ArticleLiberman DF, Kovacic SA.J Clin Microbiol. 1978 Jul;8(1):99-101.The biotypes of eight Escherichia coli K-12 strains were determined. Eight different biotypes were observed. Based upon the API biotype computer file, the appearance frequency for these K-12 biotypes ranged from 1/127 (infrequent) to 1/18,046 (rare).