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- ArticleKennedy N, Beutin L, Achtman M, Skurray R, Rahmsdorf U, Herrlich P.Nature. 1977 Dec 15;270(5638):580-5.Chimaeric plasmids carrying EcoRI fragments of the F sex factor have been used to identify proteins involved in conjugation and to assign them to tra cistrons. Most of these proteins are incorporated into the cell envelope and are individually regulated at the post-transcriptional level.