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- ArticleProteau G, Silver M.Can J Microbiol. 1977 Oct;23(10):1492-4.Cellular protein in the presence of elemental sulfur is determined by the Folin reagent after treatment with benzene. Using this procedure a generation time of 46.2 +/- 4.2 h was observed for Thiobacillus acidophilus grown on elemental sulfur.