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- ArticleArnaud P, Pandey JP, Fudenberg HH, Chapuis-Cellier C, Vittoz P, Creyssel R.Clin Genet. 1977 Aug;12(2):77-9.The validity of the Hardy-Weinberg equilibrium for Pi phenotypes was tested in a population of 1,653 randomly selected, healthy blood donors in Lyon, France. A significant departure from the equilibrium was found, with an excess of FF and SS homozygotes and a deficiency of FM heterozygotes.