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- ArticleRamachandran N, Ghiron CA.Biochim Biophys Acta. 1978 Feb 15;532(2):286-93.Evidence that indole to phenolate energy transfer causes the decrease in beta-trypsin fluorescence with increasing alkalinity is obtained indirectly from viscosity, lifetime, wavelength dependence and chemical modification studies which either exclude or minimize the involvement of other possible mechanisms.