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- ArticleJarrett WF, McNeil PE, Grimshaw WT, Selman IE, McIntyre WI.Nature. 1978 Jul 20;274(5668):215-7.Cattle in upland areas of Scotland and northern England are substantially more prone to alimentary cancer than those of the immediately neighbouring lowlands, and epidemiological evidence implicates a combination of papilloma virus and bracken in the aetiology of the disease. Here Professor Jarrett outlines the circumstantial case against these agents and discusses its implications.