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- ArticleTashiro F, Hiral K, Ueno Y.Appl Environ Microbiol. 1979 Aug;38(2):191-6.Fourteen mycotoxins were tested for inhibitory effects on ribonucleic acid polymerase of rat liver and Escherichia coli and nuclear ribonuclease H of rat liver and Tetrahymena pyriformis. These enzymes were strongly inhibited by (-)-luteoskyrin, (+)-rugulosin, patulin, and PR toxin.