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- ArticleFurano AV.Proc Natl Acad Sci U S A. 1978 Jul;75(7):3104-8.Radioactive tuf mRNA was used to detect the tuf gene in bacterial DNA that had been digested by various restriction endonucleases. Both the K-12 and the B strains of Escherichia coli contain two tuf genes, but no more than two. Salmonella typhimurium also contains duplicate tuf genes.